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Davies was born Jeremy Davies Boring in Traverse City, Michigan. The son of children`s author Mel Boring ("Davies" is his mother`s maiden name).[1] He has three siblings: Josh, the oldest, is an Air Force pilot; Zachary is his younger brother; his sister Katy is the youngest in the family.[2] His parents separated when he was young, leaving Davies to relocate to Kansas with his mother until the mid-1970s, when she passed away due to lupus, Davies then crossed the country to live with his father and stepmother in Santa Barbara, California before moving back to the Midwest in 1986 (Rockford, Iowa), where he completed high school.

Davies`s first role of note came as a bit part on a 1990 episode of Singer & Sons. In 1992 he performed on two episodes of The Wonder Years. He also appeared in small roles in the NBC TV movie Shoot First: A Cop`s Vengeance and, as a "Scruffy Kid", in the pilot for a colonial-era sitcom called 1775. Among the few substantial roles he had during this time were a youth in the Showtime thriller Guncrazy and a guest appearance on Melrose Place.

In 1993, Davies was cast in a TV commercial for Subaru in which his character compares the car to punk rock. Numerous casting directors and industry forces noticed the commercial, and soon after Davies found himself being sent feature film scripts. He began building a repertoire with films such as Twister. In 1998 he landed a pivotal role in Steven Spielberg`s Saving Private Ryan as Timothy E. Upham, an American GI linguist in Normandy, France, recruited just after D-Day by Captain John Miller (Tom Hanks) to be the interpreter on a dangerous mission to rescue a paratrooper (Matt Damon). Davies`s performance was well-received, and he has gone on to star in some 15 films, including Secretary with Maggie Gyllenhaal and James Spader, and Solaris with George Clooney.

Davies joined ABC`s hit series Lost for its fourth season in 2008. Davies plays Daniel Faraday, a physicist who parachutes onto the island as part of an extraction team hired by Charles Widmore.