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Mosley was born in Arlington, Texas, to a single mother of six children. The family was poor, and when ends did not meet in one place, they had to move to another. Between moving and fights with her mother, what Lacey describes as "a war zone going on at home", she began dabbling in drugs. She started to take drugs at the age of ten. The fighting became so intense that she was forced to move in with her grandparents in Gulfport, Mississippi. After leaving her family and using drugs, Lacey experienced what she described as a "nervous breakdown", cutting off her hair and beginning to consider suicide. Her grandmother became distressed and urged Lacey to attend church. Eventually she became a Christian. Throughout her earlier years, she was an outspoken atheist, with a pronounced talent for singing. While attending high school in Gulfport, she was the lead singer for the band "Sofa Kingdom" which frequently played at WoJo`s Coffee Shop in downtown Gulfport.
In 2000, Mosley began to play with drummer James Culpepper. The pair worked with guitarists Sameer Bhattacharya and Jared Hartmann, forming a group named Passerby, renamed Flyleaf in June 2004. She has also provided background vocals on "Run to You" and "Born Again", two tracks on Christian rock band Third Day`s 2008 album Revelation.
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