Harry Waris dating history

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Harry Ilmari Waris (until 1935 Warén; born 5 May 1893 in Saarijärvi; died 22 November 1973 in Helsinki) was a Finnish botanist. He worked across several branches of botany, with major contributions to lichenology, phycology, and developmental plant physiology. His 1920 thesis on lichen gonidia (the symbiotic algal cells within lichens) was among the first to use axenic culture for lichen algae, detailing methods for isolation, cultivation, and nitrogen utilisation. His work with the desmid genus Micrasterias informed studies of cell morphology, including the roles of the nucleus and cytoplasm in shaping developing cells. In 1953 he devised an inorganic nutrient medium (often called Waris's MY medium) for desmids, which remains in use in physiological and experimental phycology. He also reported early observations of somatic embryogenesis in Oenanthe aquatica, work that preceded later studies in the field and is part of the history of developmental plant physiology.